212 Species in Family Bythitidae
Nominal species list for Bythitidae
Scientific Name Author English Name
Acarobythites larsonae Machida, 2000 Larson's cusk
Alionematichthys ceylonensis Møller & Schwarzhans, 2008
Alionematichthys crassiceps Møller & Schwarzhans, 2008 Bighead viviparous brotula
Alionematichthys minyomma (Sedor & Cohen, 1987)
Alionematichthys phuketensis Møller & Schwarzhans, 2008 Phuket viviparous brotula
Alionematichthys piger (Alcock, 1890) Whiskered viviparous brotula
Alionematichthys plicatosurculus Møller & Schwarzhans, 2008 Folded viviparous brotula
Alionematichthys riukiuensis (Aoyagi, 1954) Bigeye cusk
Alionematichthys samoaensis Møller & Schwarzhans, 2008
Alionematichthys shinoharai Møller & Schwarzhans, 2008
Alionematichthys suluensis Møller & Schwarzhans, 2008 Sulu viviparous brotula
Alionematichthys winterbottomi Møller & Schwarzhans, 2008
Anacanthobythites platycephalus Anderson, 2008
Anacanthobythites tasmaniensis Anderson, 2008
Beaglichthys bleekeri Schwarzhans & Møller, 2007 Bleeker's viviparous brotula
Beaglichthys larsonae Schwarzhans & Møller, 2007
Beaglichthys macrophthalmus Machida, 1993 Beagle cusk
Bellottia apoda Giglioli, 1883
Bellottia armiger (Smith & Radcliffe, 1913)
Bellottia cryptica Nielsen, Ross & Cohen, 2009
Bellottia galatheae Nielsen & Møller, 2008
Bellottia robusta Nielsen, Ross & Cohen, 2009
Bidenichthys beeblebroxi Paulin, 1995
Bidenichthys capensis Barnard, 1934 Freetail brotula
Bidenichthys consobrinus (Hutton, 1876) Grey brotula
Brosmodorsalis persicinus Paulin & Roberts, 1989 Pink brotula
Brosmolus longicaudus Machida, 1993 Longtail cusk
Brosmophyciops pautzkei Schultz, 1960 Slimy cuskeel
Brosmophycis marginata (Ayres, 1854) Red brotula
Brotulinella taiwanensis Schwarzhans, Møller & Nielsen, 2005 Taiwan coralbrotula
Bythites fuscus Reinhardt, 1837 Arctic brotula
Bythites gerdae Nielsen & Cohen, 1973
Bythites islandicus Nielsen & Cohen, 1973
Calamopteryx goslinei Böhlke & Cohen, 1966 Longarm brotula
Calamopteryx jeb Cohen, 1973
Calamopteryx robinsorum Cohen, 1973 Teacher brotula
Cataetyx alleni (Byrne, 1906)
Cataetyx bruuni (Nielsen & Nybelin, 1963)
Cataetyx chthamalorhynchus Cohen, 1981
Cataetyx hawaiiensis Gosline, 1954
Cataetyx laticeps Koefoed, 1927
Cataetyx lepidogenys (Smith & Radcliffe, 1913)
Cataetyx messieri (Günther, 1878)
Cataetyx nielseni Balushkin & Prokofiev, 2005 Nielsen's catetyx
Cataetyx niki Cohen, 1981
Cataetyx platyrhynchus Machida, 1984
Cataetyx rubrirostris Gilbert, 1890 Rubynose brotula
Cataetyx simus Garman, 1899
Dactylosurculus gomoni Schwarzhans & Møller, 2007
Dermatopsis greenfieldi Møller & Schwarzhans, 2006 Greenfield's mudbrotula
Dermatopsis hoesei Møller & Schwarzhans, 2006 Hoese's mudbrotula
Dermatopsis joergennielseni Møller & Schwarzhans, 2006 Nielsen's mudbrotula
Dermatopsis macrodon Ogilby, 1896 Eastern yellow blindfish
Dermatopsis multiradiatus McCulloch & Waite, 1918 Slender blindfish
Dermatopsoides andersoni Møller & Schwarzhans, 2006 Anderson's mudbrotula
Dermatopsoides kasougae (Smith, 1943) Orange brotula
Dermatopsoides morrisonae Møller & Schwarzhans, 2006 Morrison's mudbrotula
Dermatopsoides talboti Cohen, 1966 Lesser orange brotula
Diancistrus alatus Schwarzhans, Møller & Nielsen, 2005 Winged coralbrotula
Diancistrus alleni Schwarzhans, Møller & Nielsen, 2005 Allen's coralbrotula
Diancistrus altidorsalis Schwarzhans, Møller & Nielsen, 2005 Humpbacked coralbrotula
Diancistrus atollorum Schwarzhans, Møller & Nielsen, 2005 Atoll coralbrotula
Diancistrus beateae Schwarzhans, Møller & Nielsen, 2005 Beate's coralbrotula
Diancistrus brevirostris Schwarzhans, Møller & Nielsen, 2005 Shortnosed coralbrotula
Diancistrus eremitus Schwarzhans, Møller & Nielsen, 2005 Lonely coralbrotula
Diancistrus erythraeus (Fowler, 1946) Red viviparous brotula
Diancistrus fijiensis Schwarzhans, Møller & Nielsen, 2005 Fiji coralbrotula
Diancistrus fuscus (Fowler, 1946) Dusky brotulid
Diancistrus jackrandalli Schwarzhans, Møller & Nielsen, 2005 Randall's coralbrotula
Diancistrus jeffjohnsoni Schwarzhans, Møller & Nielsen, 2005 Johnson's coralbrotula
Diancistrus karinae Schwarzhans, Møller & Nielsen, 2005 Karin's coralbrotula
Diancistrus katrineae Schwarzhans, Møller & Nielsen, 2005 Katrine's coralbrotula
Diancistrus leisi Schwarzhans, Møller & Nielsen, 2005 Leis' coralbrotula
Diancistrus longifilis Ogilby, 1899 Twinhook cusk
Diancistrus machidai Schwarzhans, Møller & Nielsen, 2005 Machida's coralbrotula
Diancistrus manciporus Schwarzhans, Møller & Nielsen, 2005 Few-pored coralbrotula
Diancistrus mcgroutheri Schwarzhans, Møller & Nielsen, 2005 Mcgrouther's coralbrotula
Diancistrus mennei Schwarzhans, Møller & Nielsen, 2005 Menne's coralbrotula
Diancistrus niger Schwarzhans, Møller & Nielsen, 2005 Dark coralbrotula
Diancistrus novaeguineae (Machida, 1996) New guinea viviparous brotula
Diancistrus pohnpeiensis Schwarzhans, Møller & Nielsen, 2005 Pohnpei coralbrotula
Diancistrus robustus Schwarzhans, Møller & Nielsen, 2005 Robust coralbrotula
Diancistrus springeri Schwarzhans, Møller & Nielsen, 2005 Springer's coralbrotula
Diancistrus tongaensis Schwarzhans, Møller & Nielsen, 2005 Tonga coralbrotula
Diancistrus typhlops Nielsen, Schwarzhans & Hadiaty, 2009
Diancistrus vietnamensis Schwarzhans, Møller & Nielsen, 2005 Vietnam coralbrotula
Didymothallus criniceps Schwarzhans & Møller, 2007
Didymothallus mizolepis (Günther, 1867) Smalleye cusk
Didymothallus nudigena Schwarzhans & Møller, 2011
Didymothallus pruvosti Schwarzhans & Møller, 2007
Dinematichthys iluocoeteoides Bleeker, 1855 Yellow pigmy brotula
Dinematichthys trilobatus Møller & Schwarzhans, 2008 Trilobed viviparous brotula
Diplacanthopoma alcockii Goode & Bean, 1896
Diplacanthopoma brachysoma Günther, 1887
Diplacanthopoma brunnea Smith & Radcliffe, 1913
Diplacanthopoma japonicus (Steindachner & Döderlein, 1887)
Diplacanthopoma jordani Garman, 1899
Diplacanthopoma kreffti Cohen & Nielsen, 2002 Deepbody cusk
Diplacanthopoma nigripinnis Gilchrist & von Bonde, 1924
Diplacanthopoma raniceps Alcock, 1898
Diplacanthopoma riversandersoni Alcock, 1895
Dipulus caecus Waite, 1905 Orange eelpout
Dipulus hutchinsi Møller & Schwarzhans, 2006 Hutchin's mudbrotula
Dipulus norfolkanus Machida, 1993
Ematops randalli (Cohen & Wourms, 1976) Randall's viviparous brotula
Eusurculus andamanensis Schwarzhans & Møller, 2007 Andaman viviparous brotula
Eusurculus pistillum Schwarzhans & Møller, 2007
Eusurculus pristinus Schwarzhans & Møller, 2007 Pristine viviparous brotula
Fiordichthys paxtoni (Nielsen & Cohen, 1986) Baldhead cusk
Fiordichthys slartibartfasti Paulin, 1995
Grammonus ater (Risso, 1810)
Grammonus claudei (Torre y Huerta, 1930) Reef-cave brotula
Grammonus diagrammus (Heller & Snodgrass, 1903) Purple brotula
Grammonus longhursti (Cohen, 1964)
Grammonus minutus Nielsen & Prokofiev, 2010
Grammonus nagaredai Randall & Hughes, 2008
Grammonus opisthodon Smith, 1934 Bighead brotula
Grammonus robustus Smith & Radcliffe, 1913
Grammonus thielei Nielsen & Cohen, 2004
Grammonus waikiki (Cohen, 1964)
Grammonus yunokawai Nielsen, 2007
Gunterichthys bussingi Møller, Schwarzhans & Nielsen, 2004 Bussing's mudbrotula
Gunterichthys coheni Møller, Schwarzhans & Nielsen, 2004 Cohen's mudbrotula
Gunterichthys longipenis Dawson, 1966 Gold brotula
Hastatobythites arafurensis Machida, 1997 Spinyhead cusk
Hephthocara crassiceps Smith & Radcliffe, 1913
Hephthocara simum Alcock, 1892
Lapitaichthys frickei Schwarzhans & Møller, 2007
Lucifuga dentata Poey, 1858 Toothed Cuban cusk-eel
Lucifuga inopinata Cohen & McCosker, 1998
Lucifuga lucayana Møller, Schwarzhans, Iliffe & Nielsen, 2006 Lucaya cavebrotula
Lucifuga simile Nalbant, 1981
Lucifuga spelaeotes Cohen & Robins, 1970 New Providence cusk-eel
Lucifuga subterranea Poey, 1858 Cuban cusk-eel
Lucifuga teresinarum Díaz Pérez, 1988
Majungaichthys agalegae Schwarzhans & Møller, 2011
Majungaichthys simplex Schwarzhans & Møller, 2007
Mascarenichthys heemstrai Schwarzhans & Møller, 2007
Mascarenichthys microphthalmus Schwarzhans & Møller, 2007
Mascarenichthys remotus Schwarzhans & Møller, 2011
Melodichthys hadrocephalus Nielsen & Cohen, 1986
Microbrotula andersoni Schwarzhans & Nielsen, 2011 Anderson's viviparous brotula
Microbrotula bentleyi Anderson, 2005 Many-ray cusk
Microbrotula geraldalleni Schwarzhans & Nielsen, 2012
Microbrotula greenfieldi Anderson, 2007
Microbrotula hamata Schwarzhans & Nielsen, 2011
Microbrotula punicea Anderson, 2007
Microbrotula queenslandica Anderson, 2005 Queensland cusk
Microbrotula rubra Gosline, 1953
Monothrix polylepis Ogilby, 1897 Littoral cusk
Nielsenichthys pullus Schwarzhans & Møller, 2011 Nusa penida viviparous brotula
Ogilbia boehlkei Møller, Schwarzhans & Nielsen, 2005 Boehlke's coralbrotula
Ogilbia boydwalkeri Møller, Schwarzhans & Nielsen, 2005 Professor brotula
Ogilbia cayorum Evermann & Kendall, 1898 Key brotula
Ogilbia cocoensis Møller, Schwarzhans & Nielsen, 2005 Coco coralbrotula
Ogilbia davidsmithi Møller, Schwarzhans & Nielsen, 2005 Cortez brotula
Ogilbia deroyi (Poll & van Mol, 1966) Deroy's cuskeel
Ogilbia galapagosensis (Poll & LeLeup, 1965) Galapagos cuskeel
Ogilbia jeffwilliamsi Møller, Schwarzhans & Nielsen, 2005 William's coralbrotula
Ogilbia jewettae Møller, Schwarzhans & Nielsen, 2005 Jewett's coralbrotula
Ogilbia mccoskeri Møller, Schwarzhans & Nielsen, 2005 Mccosker's coralbrotula
Ogilbia nigromarginata Møller, Schwarzhans & Nielsen, 2005 Blackmargin brotula
Ogilbia nudiceps Møller, Schwarzhans & Nielsen, 2005 Slickhead brotula
Ogilbia robertsoni Møller, Schwarzhans & Nielsen, 2005 Brown brotula
Ogilbia sabaji Møller, Schwarzhans & Nielsen, 2005 Curator brotula
Ogilbia sedorae Møller, Schwarzhans & Nielsen, 2005 Notchspine brotula
Ogilbia suarezae Møller, Schwarzhans & Nielsen, 2005 Shy brotula
Ogilbia tyleri Møller, Schwarzhans & Nielsen, 2005 Tyler's coralbrotula
Ogilbia ventralis (Gill, 1863) Gulf brotula
Ogilbichthys ferocis Møller, Schwarzhans & Nielsen, 2004 Ferocious coralbrotula
Ogilbichthys haitiensis Møller, Schwarzhans & Nielsen, 2004 Haiti coralbrotula
Ogilbichthys kakuki Møller, Schwarzhans & Nielsen, 2004 Kakuk's coralbrotula
Ogilbichthys longimanus Møller, Schwarzhans & Nielsen, 2004 Longfinned coralbrotula
Ogilbichthys microphthalmus Møller, Schwarzhans & Nielsen, 2004 Small-eyed coralbrotula
Ogilbichthys puertoricoensis Møller, Schwarzhans & Nielsen, 2004 Puerto Rico coralbrotula
Ogilbichthys tobagoensis Møller, Schwarzhans & Nielsen, 2004 Tobago coralbrotula
Paradiancistrus acutirostris Schwarzhans, Møller & Nielsen, 2005 Sharpnosed coralbrotula
Paradiancistrus christmasensis Schwarzhans & Møller, 2011 Christmas viviparous brotula
Paradiancistrus cuyoensis Schwarzhans, Møller & Nielsen, 2005 Cuyo coralbrotula
Paradiancistrus lombokensis Schwarzhans & Møller, 2007 Lombok viviparous brotula
Parasaccogaster melanomycter (Cohen, 1981)
Parasaccogaster normae (Cohen & Nielsen, 1972)
Parasaccogaster rhamphidognatha (Cohen, 1987)
Porocephalichthys dasyrhynchus (Cohen & Hutchins, 1982) Shaggy cusk
Pseudogilbia sanblasensis Møller, Schwarzhans & Nielsen, 2004 San Blas coralbrotula
Pseudonus acutus Garman, 1899
Pseudonus squamiceps (Lloyd, 1907)
Saccogaster brayae Nielsen, Schwarzhans & Cohen, 2012
Saccogaster hawaii Cohen & Nielsen, 1972
Saccogaster horrida Nielsen, Schwarzhans & Cohen, 2012
Saccogaster maculata Alcock, 1889
Saccogaster nikoliviae Nielsen, Schwarzhans & Cohen, 2012
Saccogaster parva Cohen & Nielsen, 1972
Saccogaster staigeri Cohen & Nielsen, 1972
Saccogaster tuberculata (Chan, 1966) Bagbelly cusk
Stygnobrotula latebricola Böhlke, 1957 Black brotula
Thalassobathia nelsoni Lee, 1974
Thalassobathia pelagica Cohen, 1963
Thermichthys hollisi (Cohen, Rosenblatt & Moser, 1990)
Timorichthys angustus Nielsen, Okamoto & Schwarzhans, 2013
Timorichthys disjunctus Nielsen & Schwarzhans, 2011
Tuamotuichthys bispinosus Møller, Schwarzhans & Nielsen, 2004 Two-spined slopebrotula
Tuamotuichthys marshallensis Nielsen, Schwarzhans, Møller & Randall, 2006 Marshall slopebrotula
Tuamotuichthys schwarzhansi Nielsen & Møller, 2008
Typhliasina pearsei (Hubbs, 1938) Mexican blind brotula
Ungusurculus collettei Schwarzhans & Møller, 2007 Collette's viviparous brotula
Ungusurculus komodoensis Schwarzhans & Møller, 2007 Komodo viviparous brotula
Ungusurculus philippinensis Schwarzhans & Møller, 2007 Philippines viviparous brotula
Ungusurculus riauensis Schwarzhans & Møller, 2007 Riau viviparous brotula
Ungusurculus sundaensis Schwarzhans & Møller, 2007 Sunda viviparous brotula
Ungusurculus williamsi Schwarzhans & Møller, 2007 Williams' viviparous brotula
Zephyrichthys barryi Schwarzhans & Møller, 2007
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