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| Sound produced | yes,passive & active sound production |
| Type of sound produced | swimming noise, knocks, thumps |
| Sound production organ | swim bladder & pneumatic duct |
| Sound mechanism | may be escape sounds (discharge of gas through pneumatic duct) |
| Behavioural context | swimming & handling |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | active sound production (knocks & thumps) only recorded during handling, recorded with bandpass filter: 30 -1200 Hz |